Beavers can and do dramatically change the landscape. The beaver is a keystone species-their skills as foresters and engineers create and maintain ponds and wetlands that increase biodiversity, purify water, and prevent large-scale flooding. Biologists have long studied their daily and seasonal routines, family structures, and dispersal patterns. As human development encroaches into formerly wild areas, property owners and government authorities need new, nonlethal strategies for dealing with so-called nuisance beavers. At the same time, the complex behavior of beavers intrigues visitors at parks and other wildlife viewing sites because it is relatively easy to observe.In an up-to-date, exhaustively illustrated, and comprehensive book on beaver biology and management, Dietland Muller-Schwarze gathers a wealth of scientific knowledge about both the North American and Eurasian beaver species. The Beaver is designed to satisfy the curiosity and answer the questions of anyone with an interest in these animals, from students who enjoy watching beaver ponds at nature centers to homeowners who hope to protect their landscaping. Photographs taken by the authors document every aspect of beaver behavior and biology, the variety of their constructions, and the habitats that depend on their presence. Beaver facts:*Just as individual beavers shape their immediate surroundings, so did the distribution of beavers across North America influence the paths of English and French explorers and traders. As a result of the fur trade, beavers were wiped out across large areas of the United States. Reintroduction efforts led to the widespread establishment of these resilient animals, and now they are found throughout North America, Europe, and parts of the southern hemisphere.*Beaver meadows provided early settlers with level, fertile pastures and hayfields.*Based on the fossil record, the smallest extinct beaver species were the size of a muskrat, and the largest may have reached the size of a black bear (five to six times as large as today's North American beavers). Beaver-gnawed wood has been found alongside the skeleton of a mastodon.*Some beavers remain in the home lodge for an extra year to assist their parents in raising younger siblings. They feed, groom, and guard the newborn kits.*In 1600, beaver ponds covered eleven percent of the upper Mississippi and Missouri Rivers' watershed above Thebes, Illinois. Restoring only 3 percent of the original wetlands might suffice to prevent catastrophic floods such as those in the early 1990s.
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Extrait
The Beaver
Dietland MüllerSchwarze
The Beaver Its Life and Impact, Second Edition
Comsock Pubîsîng Assocîaes, a dîvîsîon o Corne Unîversîy Press, Iaca and ondon
A rîgs reserved. Excep or brîe quoaîons în a revîew, îs book, or pars ereo, mus no be reproduced în any orm wîou permîssîon în wrîîng rom e pubîser. or înormaîon, address Corne Unîversîy Press, Sage House, 12 Eas Sae Sree, Iaca, New York 180.
he irs edîîon o îs book was pubîsed wî e aîd o a gran rom e Humane Socîey o e Unîed Saes. Ar or beaver racks used couresy o Kîm A. Cabrera.
Second edîîon pubîsed 2011 by Corne Unîversîy Press
îrs edîîon pubîsed 2003 ashe Beaver: Natural HIstory of a Wetlands EngIneer, by Dîeand Müer-Scwarze and îxîng Sun, by Corne Unîversîy Press
Prîned în e Unîed Saes o Amerîca
îbrary o Congress Caaogîng-în-Pubîcaîon Daa Müer-Scwarze, Dîeand. he beaver : îs îe and împac / Dîeand Müer-Scwarze. — 2nd ed. p. cm. Incudes bîbîograpîca reerences and îndex. ISBN 978-0-801-010- (co : ak. paper) 1. Beavers. I. Tîe. Q737.R32M8 2011 99.37—dc22 20110001
Corne Unîversîy Press srîves o use envîronmenay responsîbe suppîers and maerîas o e ues exen possîbe în e pubîsîng o îs books. Suc maerîas încude vegeabe-based, ow-VOC înks and acîd-ree papers a are recyced, oay corîne-ree, or pary composed o nonwood ibers. or urer înormaîon, vîsî our websîe a www.cornepress.corne.edu.
Co prînîng 10 9 8 7 3 2 1
Contents
Preace vîî Inroducîon xî
Part I The Organism . Now and hen: he Specîes, Incudîng ossîs . orm, Weîg, and Specîa Adapaîons . Dîvîng and hermoreguaîon: rom and Mamma o Semîaquaîc Desîgn and uncîon . Energy Budge
Part II Behavior . amîîes as Socîa Unîs . Communîcaîon by Scen and Sound . Inrasrucure: Dams, odges, Traîs, and Canas . Beaver Tîme . ood Seecîon
Part III Populations . Reproducîon, Deveopmen, and îe Expecancy . Popuaîon Densîîes and Dynamîcs . îndîng a Home: Dîspersa
PartIVEcology . Were hey îve and Wy: Habîa Requîremens . Moraîy and Predaors . Parasîes and Dîseases . Maker o andscapes: Creaîng Habîa or Pans, Anîmas, and Peope
vi |Contents
Part V Beaver and People: Conservation, Use, and Management . “Here beore Crîs”: ur Trade, e “Beaver Repubîc” (Hudson’s Bay Company), and ur Trappîng Today . Reînroducîons and Oer Transpans . “Nuîsance Beavers” Caîm heîr and . Needed: An Ecosysems Engîneer or Habîa Resoraîon and Oer Servîces . îvîng wî Beavers: Conservaîon and Proacîve Managemen
Index
Preface
e beaver pays an enormousy împoran roe în oday’s ecoogy. In e 21s T cenury, îs anîma provîdes ecosysem servîces suc as creaîng and maîn-aînîng weands, conroîng Loods, împrovîng waer quaîy, creaîng abîa or pans and anîmas, and prevenîng soî erosîon. As awetlands engIneer, e beaver buîds dams and dîgs cannes. As akeystone specIes, î susaîns ponds and weands. Bu wen spreadîng as a non-naîve o new areas—as în Sou Amerîca —î can become anInvasIve specIes, an undesîrabe raî. he wo specîes o beaver are e Nor Amerîcan beaver,Castor canadensIs, and e Eurasîan beaver,C. iber, bo o wîc are covered în îs book. Gîven my ied researc experîence în Nor Amerîca, many exampes are drawn rom Castor canadensIs. hîs anîma dazzes wî îs exensîve consrucîons, demands respec as a survîva arîs, and—wen în conLîc wî umans—caenges e bes mînds în wîdîe managemen. Many cîîzens’ groups and proessîonas srîve o resove ose conLîcs and ind ways or us o coexîs wî beavers. In recen îmes, e Nor Amerîcan beaver and e Eurasîan beaver ave posed opposîe probems or wîdîe managers. In Nor Amerîca e managers soug o îmî e damage o uman înrasrucures by burgeonîng beaver pop-uaîons. In e counrîes a make up e regîon o Eurasîa, owever, ey worked o proec beavers and buîd up vîabe popuaîons. ur rappîng as decîned în Nor Amerîca, eadîng o rapîd sauraîon o suîabe beaver abîa. As Nor Amerîcan beavers move back îno eîr ormer abîa, ey ind î deveoped and conLîcs ensue: beavers Lood roads, parkîng os, and go courses, and cu down ornamena and ores rees. he beaver be-comes a poîîca symbo, pîîng andowners agaîns anîma rîgs’ advocaes, and wîdîe managers agaîns concerned cîîzens. In Europe, în a specacuar comeback, Eurasîan beaver popuaîons ave re-bounded rom near exîncîon. hey were reduced o abou beavers one undred years ago, bu now coun abou a a mîîon, wî over one undred ousand în e sma counry o îuanîa aone. A îs appened în a deve-oped andscape, aoug îuanîa’s uman popuaîon does no ave e same
| vii
viii |Preface
densîy as esewere în Europe. Reînroducîons are conînuîng, wî e aes occurrîng în Scoand. Now bo conînens ace e same îssue: ow o ind a baance beween maînaînîng eay beaver popuaîons and mînîmîzîng e damage ey înLîc by Loodîng and ree-cuîng. hîs remarkabe specîes as come u cîrce: rom seemîngy îmîess abundance, o near exîncîon în vas areas o Nor Amer-îca vîa e reeness and ruînous ur rade and abîa cange, o a new abun-dance oday în a prooundy canged andscape. he irs edîîon o îs book (he Beaver: Natural HIstory of a Wetlands EngI-neer, ) was co-auored by Dr. îxîng Sun, one o my ormer graduae su-dens. I aîmed o i a serîous gap în e wîdîe îeraure, because a modern, compreensîve book on beaver bîoogy or coege and îg scoo sudens and e genera reader dîd no exîs a e îme. he scîenîic îeraure was scaered over a vas unîverse o academîc journas. hîs caed or a compîaîon o conem-porary înormaîon on beaver bîoogy, accessîbe o scîenîs and non-specîaîs aîke. hîs second edîîon încorporaes e aes researc. Prevîousy no seen be-avîor îs descrîbed, suc as e “sîck dîspay.” Saeîe poograpy reveaed e onges beaver dam ye dîscovered. Cîmae cange appears o împac beavers: A Yeowsone Naîona Park, e growîng season now ass our weeks onger, boosîng wîow grow, wîc în urn beneis beavers. Beaver popuaîons în paces as ar apar as Easern Europe and Tîerra de uego are rapîdy cangîng. We are gaînîng more experîence în managîng beavers or abîa împrovemen and proecîon o rare specîes. înay, recen years ave seen sopîsîcaed ge-neîc sudîes wîc benei conservaîon o beaver popuaîons. Many coeagues and assîsans ave eped o coec e orîgîna ied daa revîewed în îs book. I ank Tîm Scwender, ecnîcîan, or îs dedîcaîon. Undergraduae assîsans, parîcuary Rîcard Corradî, Susan Heckman, înda Moreouse, and Bren Speîcer spen couness ours sîîng quîey a beaver ponds, suFerîng aackîng backLîes and mosquîoes. Our work was suppored by saF a Aegany Sae Park, e ae Dîck Sage a Hunîngon Wîdîe ores în Newcomb, and arry Raman a Cranberry ake Bîoogîca Saîon în e Adîrondacks. My wîe Crîsîne Müer-Scwarze and dauger Annee Müer-Scwarze assîsed în coecîng daa în e ied. Ideas were excanged wî or-mer graduae sudens Drs. Peer Houîan, Bruce Scue, and îxîng Sun; Sa-sîa Bembenek, Meagan Boîce-Green, Heaer Brasear, Krîsen Buecî, Bre Mosîer, Rebecca Coeman Quaî, Maryann Scwoyer, and Rober G. Wes. My coeague and rîend, e ae Dr. Rober M. “Mî” Sîverseîn, coaboraed on many experîmens, joîned us în e ied and asked peneraîng quesîons. Our work was suppored în par by grans rom e U.S. Naîona Scîence oundaîon.
Preface | ix
Uncouned caers wî “beaver probems” eped us o appy e resus o our ied researc o beaver managemen. hroug conversaîons wî co-eagues, wîdîe managers, propery owners, îgway superînendens, and cîî-zens’ groups I began o înk abou ways o appy bîoogîca and ecoogîca knowedge or beer and wîser managemen.Basîc and appîed researc work and în gove. I îs my ope a e înormaîon în îs book wî ep readers o undersand e envîronmena roe o e beaver wordwîde and conrîbue o indîng ways o coexîs wî îs exraordînaryecosystems engIneer.