Plio-Pleistocene mammalian biostratigraphy of Atlantic Morocco / Biostratigraphie des mammifères Plio-Pléistocènes du Maroc atlantique. - article ; n°1 ; vol.13, pg 43-53
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Plio-Pleistocene mammalian biostratigraphy of Atlantic Morocco / Biostratigraphie des mammifères Plio-Pléistocènes du Maroc atlantique. - article ; n°1 ; vol.13, pg 43-53

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Quaternaire - Année 2002 - Volume 13 - Numéro 1 - Pages 43-53
Le récent programme de coopération franco-marocain a permis la découverte et la fouille de plusieurs riches sites dans la région de Casablanca. Le plus ancien d'entre eux est celui de Lissasfa, âgé d'au moins 5 M a., qui montre que la séquence « quaternaire » du Maroc atlantique est au moins deux fois plus longue qu'on ne le pensait naguère. Les autres sites permettent l'établissement d'un cadre biochronologique pour le Pliocène supérieur et le Quaternaire nord-africain. Ahl al Oughlam, daté d'environ 2,5 M a., est l'une des plus riches localités de tout le Cénozoique africain, mais l'Homme n'avait sans doute pas encore gagné l'Afrique du Nord à cette époque. Dans la carrière Thomas 1, le plus ancien niveau archéologique du Maroc, le niveau L, est probablement d'âge pléistocene inférieur. Dans le même ensemble de carrières, les faunes de remplissages de fissures, dont la plus ancienne est associée à l'Homo erectus de Thomas 1, peuvent être regroupées dans un même ensemble.
The recent Moroccan-French cooperation programme has brought about the discovery and excavation of several nch sites in the area of Casablanca. The earliest of these is the newly discovered locality of Lissasfa, which is at least 5 Myr old, and shows that the «Quaternary» sequence of Atlantic Morocco is at least twice as long as previously thought. The other localities provide a biochronological frame for the North-African Upper Pliocene and Quaternary Ahl al Oughlam, dated to about 2.5 Myr, is one of the richest faunal localities in the whole African Cenozoic, but man had probably not arrived in North Africa at that time. In Thomas Quarry 1, the earliest archaeological level in Morocco, level L, is probably of lower Pleistocene age. In the same complex of quarries, the faunas of cave fillings, of which the Homo erectus level of Thomas 1 looks the oldest, can be grouped within a single faunal unit.
11 pages
Source : Persée ; Ministère de la jeunesse, de l’éducation nationale et de la recherche, Direction de l’enseignement supérieur, Sous-direction des bibliothèques et de la documentation.

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Publié le 01 janvier 2002
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Denis Geraads
Plio-Pleistocene mammalian biostratigraphy of Atlantic Morocco
/ Biostratigraphie des mammifères Plio-Pléistocènes du Maroc
atlantique.
In: Quaternaire - Volume 13 - Numéro 1 - 2002. pp. 43-53.
Résumé
Le récent programme de coopération franco-marocain a permis la découverte et la fouille de plusieurs riches sites dans la région
de Casablanca. Le plus ancien d'entre eux est celui de Lissasfa, âgé d'au moins 5 M a., qui montre que la séquence «
quaternaire » du Maroc atlantique est au moins deux fois plus longue qu'on ne le pensait naguère. Les autres sites permettent
l'établissement d'un cadre biochronologique pour le Pliocène supérieur et le Quaternaire nord-africain. Ahl al Oughlam, daté
d'environ 2,5 M a., est l'une des plus riches localités de tout le Cénozoique africain, mais l'Homme n'avait sans doute pas encore
gagné l'Afrique du Nord à cette époque. Dans la carrière Thomas 1, le plus ancien niveau archéologique du Maroc, le niveau L,
est probablement d'âge pléistocene inférieur. Dans le même ensemble de carrières, les faunes de remplissages de fissures, dont
la plus ancienne est associée à l'Homo erectus de Thomas 1, peuvent être regroupées dans un même ensemble.
Abstract
The recent Moroccan-French cooperation programme has brought about the discovery and excavation of several nch sites in the
area of Casablanca. The earliest of these is the newly discovered locality of Lissasfa, which is at least 5 Myr old, and shows that
the «Quaternary» sequence of Atlantic Morocco is at least twice as long as previously thought. The other localities provide a
biochronological frame for the North-African Upper Pliocene and Quaternary Ahl al Oughlam, dated to about 2.5 Myr, is one of
the richest faunal localities in the whole African Cenozoic, but man had probably not arrived in North Africa at that time. In
Thomas Quarry 1, the earliest archaeological level in Morocco, level L, is probably of lower Pleistocene age. In the same complex
of quarries, the faunas of cave fillings, of which the Homo erectus level of Thomas 1 looks the oldest, can be grouped within a
single faunal unit.
Citer ce document / Cite this document :
Geraads Denis. Plio-Pleistocene mammalian biostratigraphy of Atlantic Morocco / Biostratigraphie des mammifères Plio-
Pléistocènes du Maroc atlantique. In: Quaternaire - Volume 13 - Numéro 1 - 2002. pp. 43-53.
doi : 10.3406/quate.2002.1702
http://www.persee.fr/web/revues/home/prescript/article/quate_1142-2904_2002_num_13_1_1702Quaternaire, 13, (1), 2002, p 43-53
PLIO-PLEISTOCENE MAMMALIAN BIOSTRATIGRAPHY
OF ATLANTIC MOROCCO
Denis GERAADS*
ABSTRACT
The recent Moroccan-French cooperation programme has brought about the discovery and excavation of several nch sites in the area of Casa
blanca. The earliest of these is the newly discovered locality of Lissasfa, which is at least 5 Myr old, and shows that the «Quaternary» sequence of
Atlantic Morocco is at least twice as long as previously thought The other localities provide a biochronological frame for the North-African Upper
Pliocene and Quaternary Ahl al Oughlam, dated to about 2.5 Myr, is one of the richest faunal localities in the whole African Cenozoic, but man had
probably not arrived in North Africa at that time In Thomas Quarry 1, the earliest archaeological level in Morocco, level L, is probably of lower
Pleistocene age In the same complex of quarries, the faunas of cave fillings, of which the Homo erectus level of Thomas 1 looks the oldest, can be
grouped within a single faunal unit.
Key-words : Morocco, Pho-Pleistocene, mammalia, biostratigraphy.
RÉSUMÉ
BIOSTRATIGRAPHIE DES MAMMIFÈRES PLIO-PLÉISTOCÈNES DU MAROC ATLANTIQUE
Le récent programme de coopération franco-marocain a permis la découverte et la fouille de plusieurs riches sites dans la région de Casablanca.
Le plus ancien d'entre eux est celui de Lissasfa, âgé d'au moins 5 M a., qui montre que la séquence «quaternaire» du Maroc atlantique est au moins
deux fois plus longue qu'on ne le pensait naguère. Les autres sites permettent l'établissement d'un cadre biochronologique pour le Pliocène supérieur et
le Quaternaire nord-africain. Ahl al Oughlam, daté d'environ 2,5 M a., est l'une des plus riches localités de tout le Cénozoique africain, mais l'Homme
n'avait sans doute pas encore gagné l'Afrique du Nord à cette époque Dans la carrière Thomas 1, le plus ancien niveau archéologique du Maroc, le
niveau L, est probablement d'âge pleistocene inférieur Dans le même ensemble de carrières, les faunes de remplissages de fissures, dont la plus ancienne
est associée à Y Homo erectus de Thomas 1, peuvent être regroupées dans un même ensemble
Mots-clés : Maroc, Pho-Pléistocène, mammifères, biostratigraphie.
site). Arambourg's synthesis (1938) dealt mainly with the INTRODUCTION
upper Pleistocene and includes many fossils of doubtful
stratigraphie provenance. Although Quaternary Vertebrate remains have been
The first serious attempt to establish a biochronology known from Morocco for more than a century, there
of Morocco and indeed, the whole Maghreb, was that has been little systematic collecting. Over a long period,
of Jaeger who systematically collected Rodents which Mammalian fossils were collected by geologists without
were studied by him (1971a & b, 1975, 1988) and Tong proper record of their exact provenance, context or asso
(1989). Unfortunately, most of Jaeger's localities lack ciations, sometimes just as curiosities for Museum dis
large Mammal remains, so his framework cannot be play. As a result, few reliable assemblages of large
extended to them. Mammal remains are available for research. Of these few
In the late sixties, following the stratigraphie studies significant samples from well-defined localities, I may
of Biberson (1961), attention focused on the Thomas cite those of Am Maarouf and Jebel Irhoud (Hominid
*UPR 2147 du CNRS, 44 rue de l'Amiral Mouchez, 75014 PARIS (and Mission Paléontologique Française au Maroc «littoral»).
Manuscrit reçu le 08/10/2001, accepté le 02/12/2001 44
Quarries south of Casablanca, where several Homo erec-
tus remains were found at that time (Ennouchi, 1969,
1972 ; Geraads & al, 1980). Renewed stratigraphie
research on these quarries was carried out from 1978
onwards by the University of Bordeaux. It is mainly
thanks to the «Programme Casablanca», initiated by
the «Institut National des Sciences de l'Archéologie et
du Patrimoine» from Rabat, in co-operation with the
«Mission Préhistorique et Paléontologique Française au
Maroc» that systematic collecting and controlled excava
tions have been undertaken in several localities around
Casablanca (map, fig. 1).
As a result, the Quaternary palaeontology of Morocco
which, except for the Rodents and the Upper Pleisto
cene, was practically unknown until quite recently, is Fig. 1 : Localisation of the main sites of the Casablanca area ment
now documented from several well-dated sites. These ioned in the text.
Fig 1 des principaux sites de la région de Casablanca sites are reviewed below, starting with the oldest. mentionnés dans le texte Upper case is used for upper teeth (Ml, M2. . .) ; lower
case is for lower teeth (ml, m2. . .).
are less reduced than those of contemporary and even ear
THE LOWER PLIOCENE lier European species, and it might belong to a lineage of
its own, long detached from the Northern group.
Lissasfa - Myocricetodon sp. This primitive Gerbillid, represen
The new locality of Lissasfa, discovered in 1995 by ted at Lissasfa by 4 teeth only, is unknown after the Mio
D. Lefevre and J.-P. Raynal, is outside the chronological cene in North Africa and Spain.
period covered by this review, since it certainly pre - Protatera davidi. This most interesting species is the
dates the Quaternary. However, it is worth referring to, most common Mammal of Lissasfa. It is again with Spa
because it demonstrates that the Plio-Pleistocene depos nish forms close in age to the Mio-Pliocene boundary
its of Casablanca certainly cover a much longer period (Alcoy, Caravaca : Agusti, 1991) that it displays the most
than previously believed. significant similarities. The genus is unknown in North
Lissasfa is a karstic filling within aeolian sandstone Africa after the late Miocene.
near Casablanca, 12 km from the coast. Up to now, it - Irhoudia sp. This is the only Ctenodactylid known in
has yielded only a few poor scrappy remains of large the Mio-Pliocene of North Africa.
Mammals, but a good collection of microfauna, consis - Atlantoxerus sp.
ting mainly of Rodents, which can provisionally be refer This unusual assemblage does not easily fit into
red to the following taxa (Geraads, 1998) : the biostratigraphic sequence reconstructed by Coiffait-
- Mus ique. A few teeth indicate the presence of this Martin (1991) in Algeria. The most similar site, unfor<

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