Taxonomy and paleobiology of a basal scincoid lizard (Lacertilia, Scincomorpha: gen. et spec. nov.) from the Messel formation (middle eocene: basal Lutetian, Geiseltalium), Germany [Elektronische Ressource] / von Sinje Weber

Sujets

• Biologie

Informations

• Publié par: goethe_universitat_frankfurt_am_main
• Publié le: 01 janvier 2001
• Nombre de lectures: 130
• Langue: English
• Poids de l'ouvrage: 46 Mo

Extrait

Taxonomy and Paleobiology of a basal Scincoid Lizard
(Lacertilia, Scincomorpha: gen. et spec. nov.)
from the Messe1 Formation Taxonomy and Paleobiology of a basal Scincoid Lizard
(Lacertilia, Scincomorpha: gen. et spec. nov.)
from the Messel Formation
(Middle Eocene: basal Lutetian, Geiseltalium), Germany
Dissertation
zur Erlangung des Doktorgrades
der Naturwissenschaften
vorgelegt beim Fachbereich Biologie
der Johann Wolfgang Goethe-Universität
in Frankfurt am Main
von
Sinje Weber
aus Frankfurt am Main
Frankfurt 2001
(DF1) vom Fachbereich Biologie der
Johann Wo1 fgang Goethe-Universität in Frankfurt als Dissertation angenommen.
Dekan: Prof. Dr. B. Streit
1. Gutachter: Dr. Ch. Winter
Prof. Dr. F.F. Steininger 2.
Datum der Disputation: 09.04.2002 Contents
I Introduction ................... ..... ............... ............................................................................ 5
1.1 Aims of this study .................................................................................................................. 5
1.2 The Messe1 Pit Fossil Site ..................................................................................................... 5
1.2.1 Geology ........................................................................................................................................... 5
1.2.2 Biostratonomy of Messe1 fossils ...................................................................................................... 6
....................................................................................... 8 1.2.3 Eocene Lake Messe1 and its surroundings
1.2.4 History of the Pit Fossil Site ............................................................................................... 9
.............................................................................................. 10 1.2.5 Previous work on Messe1 Lacertilia
2 Tawonomic and phylogenetic position of the new tavon ................... .. .................. I I
2.1 Lacertilian relationships and paleontology ....................................................................... 11
2.1.1 Main concepts in lizard taxonomy - Historical overview ............................................................. 11 14 2.1.2 Stratigraphic distribution of lizards
2.1.3 Cladistics and numerical taxonomy 14
2.1.4 Previous cladistical work using numerical taxonomy .................................................................... 17
2.2 Material and methods ......................... ..... 18
2.2.1 ......................................................................................................................................... 18
.................................................................................................................................... 2.2.2 Preparation 19
.............................................................................................................................. 2.2.3 Documentation 20
............................................................................ 21 2.2.4 Cladistical methods using numerical taxonomy
2.2.5 Terminology ................................................................................................................................. 23
2.2.6 Abbreviations ................................................................................................................................ 27
2.3 Results .................................................................................................................................. 27
2.3.1 Systematic Paleontology ................................................................................................................ 27
2.3.1 . 1 Diagnosis 28
........................................................................................................................... 2.3.1.2 Description 29
2.3.1.2.1 Specimen SMF ME 35 16, designated holotype ............................................................. 29
2.3.1.2.2 SMF ME 475a+b, paratype ........................................................... 39
2.3.1.2.3 Specimen WDC-C-MG 12211 23, designated paratype ................................................... 48
2.3.1.2.4 SMNK-PAL 38 13, ......................................................... 63
2.3.1.2.5 Specimen SMF ME 1249a+b, additional material .......................................................... 71
2.3.1.2.6 SMNK-Me 930a, .............................................................. 86
2.3.1.2.7 Cast fiom the Fossilien- und Heimatmuseum Messel, additional material ...................... 86
.................................................................................................................... 2.3.2 Intraspecific variation 98
2.3.3 Results of numerical taxonomy ..................................................................................................... 98
2.3.3.1 Cornplete matrices .................................................................................................................. 98
2.3.3.1.1 ESTES et al . (1988) ........................... .. ......................................................................... 98
2.3.3.1 . 1. 1 Matrix excluding the new genus and species ............................................................ 98
2.3.3.1.1.2 including the new and .......................................................... 101
2.3.3.1.2 EVANS & CHURE (1998) ................................................................................................ 101
2.3.3.1.2.1 Matrix excluding the new genus and species 10 1
2.3.3.1.2.2 including the new and 103
2.3.3.1.3 LEE (1998) ..................................................................................................................... 103
2.3.3.1.3.1 Matrix excluding the new genus and species 103
2.3.3.1.3.2 including the new and 103
2.3.3.1.4 CALDWELL (1999) .......................................................................................................... 105
2.3.3.1.4.1 Matrix excluding the new genus and species .......................................................... 105
2.3.3.1.4.2 including the new and 105
2.3.3.2 Matrices limited to osteological characters .......................................................................... 105
107 2.3.3.2.1 Reduced matrix of ESTES et al . (1 988) 107 2.3.3.2.1.1 Matrix excluding the new genus and species
2.3.3.2.1.2 including the new and 107
2.3.3.2.2 Reduced matrix of EVANS & CHURE (1 998) .................................................................. 109
.......................................................... 109 2.3.3.2.2.1 Matrix excluding the new genus and species
2.3.3.2.2.2 including the new and 109 2.3.3.2.3 LEE (1998) ................... .. ........................................................................................... 109
2.3.3.3 Matrix limited to characters visible in the new genus and species ....................................... 109
2.3.3.3.1 Reduced matrix of ESTES et al . (1 988) .......................................................................... 109
2.3.3.3.1.1 Matrix excluding the new genus and species .......................................................... 109
......................................................... 2.3.3.3.1.2 including the new and 111
.................................................................. 2.3.3.3.2 Reduced matrix of EVANS & CHURE (1 998) 111 111 2.3.3.3.2.1 Matrix excluding the new genus and species
2.3.3.3.2.2 including the new and 111
2.3.3.3.3 Reduced matrix of LEE (1 998) ....................................................................................... 111
2.3.3.3.3.1 Matrix excluding the new genus and species 111
2.3.3.3.3.2 including the genus and species 113
2.3.3.4 Taxa limited to Scincomorpha .............................................................................................. 113
.......................................................................... 2.3.3.4.1 Reduced matrix of ESTES et al . (1 988) 113
2.3.3.4.1.1 Matrix without characters of the new genus and species ........................................ 113
2.3.3.4.1.2 including of the new genus and ..................................... 113
.................................................................. 2.3.3.4.2 Reduced matrix of EVANS & CHURE (1998) 115
2.3.3.4.2.1 Matrix excluding the new genus and species .......................................................... 115
2.3 .3.4.2.2 including character states of the new genus and species ............................. 115
.................................................................................. 2.3.3.4.3 Reduced matrix fiom LEE (1998) 115
2.3.3.4.3.1 Matrix without character states of the new genus and species ................................ 115
2.3.3.4.3.2 including states of the new genus and 115
2.4 Discussion ........................................................................................................................... 116
2.4.1 Taxonomic position of the new genus and species based on comparative morphology .............. 116
2.4.1.1 Key characters of the new and ....................................................................... 116
2.4.1.2 Taxonomic relationship of the new genus and species to higher taxa .................................. 121
........................................................